Animalia | Arthropoda | Insecta | Coleoptera | Scarabaeidae | Macrodactylus | Macrodactylus costulatus
Eukaryota | Animalia | Arthropoda | Hexapoda | Insecta | Coleoptera | Polyphaga | Staphyliniformia | Scarabaeoidea | Scarabaeidae | Melolonthinae | Macrodactylini | Macrodactylus | Macrodactylus costulatus
COL:CS5HF | COL:N | COL:RT | COL:L2655 | COL:H6 | COL:C2L | COL:DDHNW | COL:L2PGK | COL:KVT23 | COL:6278C | COL:L2BV2 | COL:KYN | COL:CB8L6 | COL:3X4MV
(Chatton, 1925) Whittaker & Margulis, 1978 |  |  |  |  |  |  |  | Latreille, 1802 | Latreille, 1802 | MacLeay, 1819 | Kirby, 1837 | Dejean, 1821 | Bates, 1887